Johnny Joestar, born Jonathan Joestar, is a former horse racer from Danville,Kentucky, who is paralyzed from the waist down. He participates in the Steel Ball Run to follow Gyro Zeppeli in the hopes of using his Spin technique to regain use of his legs. He uses the Stand Tusk, which allows him to shoot his finger nails as bullets. As the race continues and Johnny begins to understand the Spin, Tusk evolves into different forms, termed ACTs, with differing capabilities, similarly to Koichi Hirose's Echoes.
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